M.Sc. in enzyme engineering, University of Chemical Technology in Prague
Institute of Molecular Genetics v.v.i., Academy of Sciences of the Czech Republic, Prague
Dept. Microbiology and Immunology, University of Miami School of Medicine, USA
Dept. Immunogenetics, Max-Planck-Institute for Biology in Tuebingen, Germany
Ph.D. in molecular genetics, Faculty of Science, Charles University in Prague
Organization ΦΒΔ, Miami, USA (Outstanding International Scholar)